Research and Books


Z Clay, FBM de Waal (2015) Sex and strife: post-conflict sexual contacts in bonobos. Behaviour 152 (3-4), 313-334

Z Clay, FBM de Waal (2013) Development of socio-emotional competence in bonobos. Proceedings of the National Academy of Sciences 110 (45), 18121-18126

Z Clay, FBM de Waal (2013) Bonobos respond to distress in others: consolation across the age spectrum. PLoS One 8 (1), e55206

Z Clay, K Zuberbühler (2011) The structure of bonobo copulation calls during reproductive and non‐reproductive sex. Ethology 117 (12), 1158-1169



T Gruber, Z Clay (2016) A comparison between bonobos and chimpanzees: A review and update. Evolutionary Anthropology: Issues, News, and Reviews 25 (5), 239-252

T Gruber, Z Clay, K Zuberbühler (2010) A comparison of bonobo and chimpanzee tool use: evidence for a female bias in the Pan lineage. Animal Behaviour 80 (6), 1023-1033